Alright, for those who've been following this hum hunt... First to 2deaf, the R from V3(3) to ground was 1470, and since there's a 1.5k ohm resistor in that path, it seemed good to me.
Anyhow, I rarely make more than one change at a time while on a hunt like this because I want to know exactly what fixed it. But in this case, the socket wiring was much easier to get to while the transformer wires were not in place. So I moved the reverb tranny outside of the chassis, and I did a repair to the heater wires. While looking carefully at them I realized that V3 and V2 both had inverted connections, in other words the wire from 4/5 of V4 connected to pin 9 on V3. I tried to keep them all the same, but you know, they're both green and they're twisted together.
After both changes, I fired the chassis up and... Still howled when the reverb is on and wide open... but the reverb was also NOT working. Trouble shooting with an ohmmeter, I discovered that when I unsoldered the reverb tranny red wired, the wire underneath the board had come unsoldered and dropped out. I twisted them together, and... The reverb is working, and although there is a noticeable hum when it's turned all the way up, there is no howl when it is switched on. I'm okay with it this way, not sure which move was the coup de grace, but the chassis is going back in tomorrow so I can enjoy the amp. Thank you to everyone that helped me through this, I hope I can help you one day.
-MadMax-