The mil-spec sheet says ionization is 165V, or 330V for two in series.
The REDD plan cited calls for 370V supply, 366V at the cold 0A2s. So it "should" fire both. However with the 220K, the bottom 0A2 feels the full 366V and ionizes for-sure.
When bottom 0A2 ionizes, assuming it breaks near 150V, the voltage across the 220K and the top 0A2 is about 200V.
Once you start to ionize, you want to bring the current up. Action below 5mA is undefined and wonky. Ideally we'd come up to 5mA, suggesting a 40K resistor. However after top 0A2 fires this becomes a 4mA leak around the top 0A2. Since we only have a 5mA-30mA range, now the string must pass 9mA-30mA. That does not leave much margin for E88CC variations or "370V" variation. 1mA initioal ionization is pretty solid, not to meet 144V-153V spec but enough to ensure "about 150V" drop so the upper 0A2 is sure to get enough voltage.
I don't think it HAS to be this way. You could lose the 220K and 99+% of units would fire; if not, you should replace the oA2s because they are maybe-marginal.
I don't like gas-tube hiss fed direct to amplifier stages, I don't think the circuit "needs" regulation. However it is a Holy Grail circuit so I guess you can't tamper.