I'm building an amp based on the 5D8 tweed twin (cathode biased 6L6 pair) and I'm having some trouble with very harsh distortion. I built the amp single input, single channel with octals in V1 and V2. Here's the original 5D8 schematic:

And here's how I currently have it wired:

Not pictured: a 5U4 in place of the two 5Y3's.
I'm experiencing some harsh, fizzy distortion starting at about 3 on the volume knob. The harder I dig into the strings, the harder it distorts. Playing gently yields a clean tone. Anything past noon on the volume sounds like a fuzz pedal. I'm thinking it's originating in the preamp, as changing the 6SL7's to 6SN7's dramatically reduces the volume, but the distortion onset and characteristics remain the same. Same goes for PI tube substitutions.
Originally, I didn't have the 270k to ground on V2's grid. Adding it reduced the distortion a bit.
My voltages in the preamp seemed low (a little less than 100v on V1's plate and less than 1V on V1's cathode). I dropped the B+ resistors from 10K to 4.7K, which upped the voltages a bit and slightly delayed the onset of distortion to about 5 on the volume.
The most dramatic change has been experimenting with plate resistor values. Lowering V1's plate resistor to about 50k dramatically increased the clean headroom - same goes for the 100k on V2's plates. But the distortion is still there, especially with heavy pick attack.
Other info: 6L6's biased at about 80% dissipation with a 220 ohm cathode resistor. Output transformer primary is about 9K with an 8 ohm load.
I'm having a hard time wrapping my head around what's happening in the amp, what type of distortion I'm experiencing, and how to go about fixing it. Any insight would be very much appreciated.
-Andrew