As per given advice, I have changed the bias on the power tubes with very good results. I'm now outputting 22 volts/pp to the speaker and the over all character of the amp is richer and more complex. Chiming cleans and very smooth overdrive. Thanks for that pointer.
My total plate dissipation is 125.4 mA so just under 20W per tube.
It does have some issues though, the causes of which I'm supposing are mainly due to the nature of my prototyping set up, i.e. lead dress and quality of connections. Neither of which are too terribly bad and I've taken measures previously that have actually made the circuit quite quiet as far as hum or hiss, but what I'm hearing in the background of the guitar noise is a ghosting distortion. It is distinctly not attached to the actual tone of the instrument but seems to follow it or exist parallel to it.
Also, when I removed the local negative feedback from the second triode of the 12AX7, all hell breaks loose. Many non guitar noises appear which are mostly but not exclusively tied to the volume and control pots. My input and runs to and from the tone/vol circuit are shielded and tied to ground at one end of each run. I'll be honest, I don't know what oscillation sounds like, but I believe if I did, I'd know this was it. I put the resistor and cap back, thinking maybe I knocked something askew when removing them initially and the offending sounds are gone, minus the strange background distortion tagging along.
Any thoughts about the possible causes for these symptoms? I was thinking that there might be phase issues, but that is a very uneducated assumption.
Thanks much for the pointers so far. I'm very happy with the tone I'm getting and I'm going to do some basic housekeeping and a few tube swaps as trouble shooting measures.