Gary,
It is a wonderful sounding amp when working right, so don't give up! You'll get it working OK.
The voltages on the 6L6 screen don't look right compared to the plate. Too much of a difference between the screen and plate. Should be just a few volts difference not a 100+ voltages difference. Check the screen resistor values for sure.
474v on plates and 217v on the screens is waaaayyyy off.
Have you tried other tubes? Could be a bad tube?
Other voltages look reasonable.
While you're "not" suppose to hear sound with the OT hookup backwards, I have before. I got a low volume distorted tone that would eventually oscillate & squeal as I turned up the volume. Either disconnect the negative feedback or try switching the OT wires to see if the issue is resolved.
When you are doing the trouble shooting, work with JUST the clean channel until it sounds right. It is quite possible that the OD channel will sound fine also after the clean channel sounds OK.
Be sure to safely "chopstick" all the soldering.
IF you are using a relay, I'd try safely jumpering (bypassing the relay) from the clean to the LTPI.
IF you used a passive effects loop, try bypassing that. I've had the amp do what you describe once and found out it was a bad shorting jack.
Just some things to try. PLEASE report back after trying things and let us know what you have found out.
With respect, Tubenit