There is no way that HV CT was ever soldered at the factory to pin#1 passing through the 220K grid resistor to pin #5.
Sure it was. Pin 1 was often connected to ground back in those days. Heck, the old metal 6V6s had the metal shell tied to ground. Look at this pic...

My point is that pin 1 was a ground tie point. It was convenient to connect the center tap to that ground point. It was also convenient to connect the 220K grid resistor to that same ground point. So quit thinking about the HV CT was passing through the 220K grid resistor to pin #5. That's ass backwards!
Gibson did not rely on that painted chassis to provide a good ground. That's why they strung a ground wire around the circuits. The real problem occurred when you replaced the caps but began connecting them directly to chassis (which ain't a good ground in this amp). That's why I wanted you to use gator clip leads to connect the negative end of each filter cap directly to the HV CT.
There are several ways to permanently fix this problem. Personally, I'd connect the HV CT back to pin 1. Then I'd run a wire from the negative lead of each filter cap you replaced back to the CT, or that bare wire on the pots. However you decide to do it, when all is done make sure that 220K resistor between pins 5 and 1 still measures zero ohms to the CT. That CT is like the negative terminal of a car battery. It is the ultimate ground point and everything on the schematic that should be connected to ground better have a good connection back to the CT.
Edit and fixed... Don't know why I kept saying pin 6 when I meant pin 1.