I looked the tone stack placed between the plate of v2a and the grid of v2b, and found it close to the "E series" tone stack found in the Duncan Amplification.
They are more-different than alike.
The circuit at the plate of V2A is a feedback loop to the grid of V2A (which reduces gain of that stage). Inside that feedback loop is a twin-T network, and if the junction of C5 & R9 went to ground instead of the Treble Expander pot (and R44), then that network would provide theoretically infinite attenuation at one frequency. The relative values of the twin-T components (R7, R8, R9, C5, C6, C7) determine how narrow that notch is.
Because the "infinite reduction" twin-T is sitting in a negative feedback loop, it's really infinite reduction of negative feedback at that frequency (and a band around the center frequency). Which means boosting of apparent gain at the center frequency, and the band of frequencies around it (again, the twin-T values determine the narrowness of the notch, or the width of the band that gets boosted). There is resistance between the Expander pot (and the resistance of the pot itself) to allow some variation between a minimum-boost and a maximum boost; max boost should occur when the resistance to ground is the least.
PRR's 1-resistor change sets a fixed amount of feedback for all frequencies, and can be adjusted by raising/lowering the value of the resistor jumpering the twin-T network.
Repeat all that for the Bass Expander section. The Presence control is just another way to cut the feedback around V2B to boost extreme highs.
The Treble and Bass controls between V2A and V2B (C10-C13, R11,
both R13's and the pots) are just a common James tone control. Max midrange happens when you turn the Treble and Bass pots to 0; minimum midrange happens when you turn the Treble & Bass pots full-up. In this circuit, midrange is essentially fixed at one level.