I don't think there's much practical difference between a 475 volt electrolytic and a 500 volt. However, I'd get on rated for 500 volts if it's available myself. Depends partly on what rectifier tube you've got in there. If you're using a solid-state substitute, 500 volts might not be enough safety cushion.
Is your amp the AA964 circuit or the AB1270 circuit? I'm asking because the AA964 circuit only has 3 filter caps on the power rail AFAIK. Check out Hoffman's schematic page for two versions of the AA964:
http://www.el34world.com/charts/Schematics2.phpYou'll notice that the "reservoir cap" (first in power rail - supplies OT) is 40uf in the slightly later version.
Hoffman sells a JJ four section cap can that has 40-20-20-20 sections and is rated at 500 volts that is cheaper than the AES 4x20uf cap I can see. That might be a better alternative if your amp has 4 power rail filter caps.
BTW those reddish brown coupling caps on the board look a bit worse for wear & tear, especially the righthand one that looks like it's partly melted.
Also, is the "Ground" switch still wired with the "death cap" or does it serve another purpose? I think I see wiring for a 3-prong cord there. If so, please check and make sure it's wired properly. I worked on a Blackface Princeton Reverb last year that had a very nasty habit of shocking the guitarist (or amp tech).
Hope this helps,
Chip