Your voltages appear to be okay on B+, the one I worked on recently measured 466v w/ all tubes in and bias way down. It slightly lowered after correcting bias to -39mA. You for sure have to properly spec'd PT right?
Have you carefully checked all of the contacts on all of the boards especially the main board w/ all of the tubes' sockets? I've found many suspect and outright bad contacts that all needed to be re-flowed. I resolder all tubes' sockets whether they need it or not because they mostly all do regardless how they look to you. Those boards are so cheaply made and manufactured. But that's what helps keep techs in business. Remove and check the IEC/B+ board. I found hideous cracks on solder joints there too. While you're at it, solder a 220k/1W or higher "soft start" resistor on that board to the wires that put it across the standby switch. (see pic)
Test the 100K discolored resistor and replace if necessary, maybe w/ a 1w if needed? Get the amp up and running and correctly biased. Then check your B+ voltages again w/ the chassis on a cradle or I'll turn it upside down on the head enclosure. Don't forget to put a load or speaker on the output jack (sorry, but had to say it). I'd leave it running checking for signs of excessive heat anywhere and testing what I could. Then set the chassis on a speaker cab and play it for an hour or more while monitoring everything. Sometimes it's not obvious and certain parts begin to break down after heating up more while under load. Re-check the bias afterwards also to make sure it isn't drifting.
(oops , forgot pic)