It could be nothing at all.
most tube amps sound differnt. There are the exceptional and unique ones. even built by the same mass manufacuturer on the same day with the same tubes, any two should sound different. Lead dress, layout, bias, parts tolerance,tubes, knob settings any could account for the percieved difference, more than likley the sum of all that plus some more mojo thrown in.
You said yourself it has the same voicing, just a different responce.
This is where an amp builders skills REALLY shine, tailoring the amp to get a specific "tone".
IF the amp is within tolerance, AND you know the end "tone" or "feel" or whatever you want, The guys here can help you narrow down what changes you may want to make. It was suggested to look into the NFB circuit. You already have a pot there, have you just set it according to what you did on the last amp? or have you fiddled with it yet?
LOL any how what do I know

I'm dumber than most and brave enough to prove it on a regular basis

I did notice one thing, you didn't mention that you didn't like it, that sound OK to me!
Has your friend played it? what's his opinion? I know I grade my projects on a much tougher curve than I do others, I'm a little more sensitive to nuances, more critical to minor things. a second trusted opinion may change your hearing a bit.
Ray