OK, if you've checked for m-boating, fine. I had to bring it up. Now you know what it is, so at least you've learned something. Generally---it is not selective---the circuit will oscillate obnoxiously, all the time.
Just thinking about this now, you fixed the m-boating by reversing the SPEAKER leads. OK, theoretically that achieves the same phase reversal as flipping the primary leads...BUT I would do this by reversing the primary, HV-to-tube-plate leads. Perchance...is there an internal ground connection inside your output tranny?
[drive-by suggestions follow]
I lean towards lead dress, then, along w/tubeswell, assuming you've done everything else wire-for-wire. Get your wires (except for the up-in-the-air-heater wires if that's what you have) close to the sheet metal of the chassis. Pay attention to any wires that go to the preamp tubes, pins 2 and 7. Those are your grids, those are what's gonna be sensitive.
By the way...1968 is when lots of Fenders went from the older type of wiring to the newer type. Older = green heater wire up in the air; piushback insulation, yellow wires from the board to the tube sockets. Newer = "spaghetti" wiring with plastic-insulated wires coiled around bundles of other wires like a hangman's noose. Which do you have?
Is the power transformer supposed to be a "drop-in" replacement?
Pull the tremolo tube. Still squeal?