PI to power tubes:
LARMAR or not, the R2 (lower half) of those 250K pots are parallel with the 2.2M's, so the 2.2M's don't influence grid-leak that much. At full-tilt, you are a hair under 250K. At 50%, you are two hairs under 125K. (so I don't think the 2.2M's hurt anything).
however, I thought LARMAR was designed for fixed bias amps. I don't think it does anything for you here. and,, I think this general area of the circuit is where your bass response is disappearing.
the low frequency response of the PI to output stage is determined by the 47nf coupling cap, the 250K grid leak, and the output impedance of the PI (50K?).
edit --- I was wrong on the following (now strike-thru), the 47nf always sees 250K resistance to ground to the right. bass response shouldn't be effected by wiper position.
as you turn down the MV 250K, your bass-roll off frequency climbs. So if you tried the amp with the MV set low, you might have lost some/lots of your bass response. turned up to "11", you more/less have the low freq. roll off factors of a Marshall, but as you turn down the MV,, that 250K goes south,, and bass response disappears.
preamp:
I'm with sluckey on adding a 470K or a 1M above your 1M volume. However, you can approximate the effect of such a mod by testing the volume at 50% without soldering.
usually, in a design like this, the 2nd gain stage is there to make up gain for a complex tone stack that drops the Vrms of the signal so low that it needs recovery. Where there are simple low-load volume/tone setups (like yours), the 2nd stage isn't used, signal is big enough to go to PI. additionally, those 2 stage preamps usually have a 2nd channel that gets mixed with another channel (and thus attenuate everything). PLUS the gain-increasing cathode bypass caps are usually on one gain stage or the other, not both. PLUS no neg. feedback. You don't have any of these components that reduce gain in other circuits.
So, yea "too much gain".. you built a gain monster!!
Is there any combination of the volume controls that yield a decent sound?