Adam,
You've done so much right already with this amp, it is truly puzzling why you aren't getting a strong OD tone.

On reply #25 you said:
Today I followed your Trouble shooting #4.OD sound appered ,but with huge OD signal.It is too high, because after a moment of pure OD signal , cranch and clatter appers.
That is an indicator to me of a bad connection or a short or faulty part.
Here is the order of what I personally try:
1) safely & vigorously chopstick around the OD section & include the OD pots, OD sockets, and relay. Anything crackle
or do you get noise increase? IF you have a volume jump, then there is a short or something.
2) check voltages when switching relay. Does anything change? (Your reported voltages look OK to me)
3) with amp turned off and power drained, I'd gently pull on all the wire leads of resistors and caps in the OD section.
Do any wires slide from inside a cap or resistor? Maybe there is a broken wire inside the resistor or capacitor body?
4) I'd disconnect each shielded wire at each OD pot (trim, drive, level) and test continuity to make sure your signal wire
isn't shorted to ground
Also check continuity with the shielded wire signal path on both ends of the shielded wire. I have had several
occasions that the fragile signal wire inside the shielded wire was broken. You can use your meter to check this
and you can safely bypass the shielded wire by paralleling an insulated alligator clipped wire in parallel to the
shielded wire. Yes, that would be noisey BUT if you had a huge volume increase, it would indicate that you
need to replace that shielded wire as it had a short in it.
5) While you have the shielded wire disconnected on each pot, you might consider creating a "test" to see if the pot is
bad in some regard. I have had a couple of pots go bad on me and not allow signal to go thru. See below. What you
are doing is bypassing each pot to see if that pot was the problem?
6) Try all the different 5751, 12AT7, 12AY7 tubes ............. etc ...........
7) completely disconnect the 470k & 250p from the signal path to the OD trim pot
8) Completely remove relay switching from the chassis & temporarily hardwire clean to OD to LTPI
I would try everything possible prior to removing the main board. Your wiring is so neat, that I hate to see you remove the main board.
With respect, Tubenit