Could you post some hi-rez pics of this amp? Are the power tubes red plating?
The resistance from pin 3 to ground is 55.8 ohms which is my cathode bias resistance.
You actually measured that with one meter lead connected directly to pin 3 and the other meter lead connected to chassis?
I keep harping about the cathode pin 3 because there's a confusing conflict with the voltages you posted in the original message. The plate voltage is dropping extremely low which indicates that a very high current is flowing through the tubes. BUT, you only have 3.5v on the cathode and that indicates there is very little current flowing through the tubes. This would make sense ***IF*** the cathode resistor was very small. This is the easy part to troubleshoot. But if the cathode resistance really is 56Ω, we need to look elsewhere.
Start by measuring the voltage of pin 2 for all four output tubes. They should all be zero volts. Next, measure the resistance of pin 2 (meter lead directly on the socket pin) of all tubes to chassis. Should be about 222K for all four tubes.
Now check the resistance of the OT primary. Each side to center tap will likely be a couple hundred ohms and should be about the same. A very high reading is not good and could cause the symptoms you describe.
If the cathode and grid measurements and OT resistance are correct we may need to look at the power supply or a bad connection between the power supply and the tube plates.
Do these tests and report back. Then we'll dive deeper if we need to.