...I've been working on a '78 TR135 with the same hum issues. It had some weird breakup that was fixed with new power tubes...
What idle plate or cathode current are each of the power tubes drawing?
...tried pulling power tubes like was suggested to OP...
What happened to the hum?
How about with the power tubes in place but the tube for phase splitter V6 removed?
...Maybe all the haphazard wiring is to blame? Or the layout in general? The balance/bias circuit does goes across the amp to get to its spot on the rear of the chassis, where the wires to the grids go around/across a couple tubes. Blackface bias layout didn't get near as much signal.
I've changed the balance pot to real bias pot, and it has made no difference in terms of hum. The original balance circuit added hum if anything...
The power tube matching / bias balance control is pretty much a necessity with this range of amps, due to the combination of the distributed load OT and poorly filtered HT. Normally with UL / DL power amp arrangements, the HT to the OT is well filtered with a CLC supply, eg
http://schems.com/schematicheaven.net/manu/sunn/sunn_model_t.pdfIt's all down to the screen grids; the clue is in the name, these are grids and have gain

so any HT ripple fed to them will modulate plate current.
I doubt many tube vendors include g2 gain in their tube matching characteristics, so failing that the bias needs tweaking to get those ripple currents to match on both p-p sides of the OT and so cancel out at idle.
This is much less of an issue with a regular pentode p-p output arrangement, as the HT supply to the screen grids is almost always much better filtered.
The wiring to the bias balance control may have looked messy but it's unlikely to have caused any performance issues.
...Bober also says this about the balance circuit; https://www.premierguitar.com/articles/22353-ask-amp-man-restoring-a-1979-fender-twin-reverb ...
Although he's a great tech, I think Jeff Bober is off target with some aspects of his analysis above.
1/ I think Fender introduced bias balance as early as 1965 with the AB165 Bassman; that's still in the golden age of tube manufacture (indeed, when did vintage tube manufacture ever go bad as he suggests?). Tubes have always had wide spread of acceptable performance characteristics either side of their type bogie, eg see p2 of
http://www.mif.pg.gda.pl/homepages/frank/sheets/084/k/KT88.pdf re the recommendation for a bias voltage range of +/-25%.
So to have reasonably similar power tube idle currents for p-p amps, the options have always been to buy selected power tubes, select suitable sets in-house, or to have independent bias or bias balance.
2/ With Fender, the 'hum balance' control was always in the heater circuit.
3/ The 'power tube matching' control function is still beneficial even with apparently 'matched' tube sets, as the tubes' g2 characteristics are not generally included in the matching criteria.
...I've changed the balance pot to real bias pot, and it has made no difference in terms of hum. The original balance circuit added hum if anything...
For the reasons noted above, I suggest to re-instate the 'power tube matching' control, and to fit an additional trimmer inside to set the overall bias voltage level.
... I put my meter on the center tap and find not just 250VDC (to be expected) but 545VAC
Is that normal?...
Yes, I think that's normal, the issue is that your meter can't provide Vac measurements when there's Vdc present.
A better meter, eg Fluke 87, should give reliable measurements in such circumstances.