Sorry for the side-track.
> The reverb driver is 1/2 12AX7 (not 12AT7).
2 things smell wrong.
* 12AT7 will bias-up at 8V; 12AX7 nearer 2V. So the grid of the 12AX7 will start distorting stuff 4 times sooner than 12AT7.
* The 220k are for mixing the two channels. If you are switching instead, the 220k should be shorts.
It may be *both* items. The 12AX7 grid distorts sooner, but it might not matter so much if fed directly from a preceding stage; but the added 220k allows 12AX7 grid distortion to screw-up the main-path signal more.
I am wondering why 12AX7 for what is really a heavier load than an AX should handle. Can you put a 12AT7 in there?
If you insist on one tube-type throughout, try one of the 220k you took out directly at the grid of the 12AX7 reverb driver (between 500p+1Meg and the AX grid) so when it overloads its input goes to 221k not 1k. Without the now-pointless 220k mix-resistors, so the signal comes from the 39k of V1 or V2, an added 221k load is small flavor not fart.
A more complicated option is to change the 1Meg at driver grid to a 1Meg "Drive" pot, wiper to grid. Logic suggests 12AX7 needs 4 times -less- drive than 12AT7, so you turn-down considerably. The 12AX7 grid overloads later, and the top-of-pot resistance isolates this distortion from the main path. This might not be a permanent panel control-- you might find that setting it at 30% from the bottom works great, and replace it with a 680k:330k divider.