My limited understanding and experimenting has shown that keeping to the plan is better for time saving and blown parts, but the mistakes also give insight.
My hunch is you have to have that 3rd 'reservoir' cap to maintain an even voltage as the input signal voltage swings or you get 'bad sounds'. The cap will boost to cover the drain smoothing the output signal..I think. But the EE's here will explain if they have time, but most of this knowledge is book learnin' per internet, and our own investment in pondering and doing what your doing now. Trial and error, but be sure to be safe - that fuse ought to be covered of course, but you know how your using this.
'Reservoir' is a good visualization for what is happening, ie a plumbing system. But it also is a scaled proportional system with relativities involved-Ithink

. Like some tweed designs running 2 or more preamp tubes off 1 node with a small value cap with no problem. This I assume works because the PT has enough umph to push for demand all the way at the other end. Because copper is expensive now it seems MAYBE modern designs get by with smaller TX using now cheap caps at higher reservoir capacity to supply the umph needed...again I'm guessing.

You mention this PT being small, so this lends to not having enough voltage 'pressure' down stream...I think/guess.
I'm not an expert for sure and have no over shoulder mentor. Have tinkered with many builds now, and PRR's council is learned so either tag on another cap and R over the 2 already or get a more potent PT...? Anyway is a interesting hobby that I have found to deepen my insights into other areas understanding modern auto electronic diagnostics, it is fun now to learn that where I assumed before I could never do it.
Oh another thought The TX's are awefully close maybe that could be doing something...or not. Good luck, The little 2-au7 has always been intriqueing.