The caps cause trouble for meter reading.
DC Operation. An ohm meter reads resistance by putting a small DC voltage across the resistor(s) under test. To simplify I have drawn the Tonesatck as a fixed shunt resistor to ground; and the MV pot, turned to full resistance as a fixed 1M resistor to ground. So we have, for simplicity's sake, 2 shunt resistors in parallel. The Tonestack is really more complex -- it contains several resistors and caps in filter circuits. The end result is a frequency-selective shunt to ground. But DC has "no frequency", and it is blocked by the filter caps in the Tonestack. This screws with an ohm meter's reading.
And, the MV pot is probably also in a parallel with the grid leak resistor of the next tube stage. If you disconnect the wires from lugs 1 & 2 of the MV pot, & leave lug 3 connected to ground (or not), you should get a proper DC ohm's reading across the MV pot which matches its stated value, because it won't be in parallel with any other resistors anymore.
AC Operation. Signal is AC which will pass through the RC filters of the Tonestack with certain frequency attenuation. Any particular RC filter will act as as a certain fixed resistance to any particular frequency. But a simple ohm meter reading does not measure this, because the filter caps in the circuit block its reading.