I'm agreeing with sluckey on this. I also do not see anything in common with the linked tilt tone controls.
To me, this is an artifact of drawing. Let's talk in terms of sides of the wheels, but I'll describe thinkning about it differently. If I was a MS Paint wizard, I'd draw this up in a hurry.
Go around the "top" of the wheel. You have series 150k and 680k resistors, and a 0.0033uF cap to ground. Think of this as a "T" instead, with the series resistors forming the top of the T and the cap to ground forming the vertical stem of the T. Then you'll recognize this as a T-filter, which shunts highs to ground; it's the low-pass part of the tone circuit.
Go around the "bottom" of the wheel. You have series 0.0033uF and 500pF caps, and a 68k resistor to ground. Thinking as a T again, the caps form the top of the T, and the resistor to ground forms the stem. This T is flipped upside down for convenience of drawing. This T-filter is a high pass, and tends to shunt lows to ground.
An alternate way to view the arrangment is a pair of pi-filters, with 2 series elements, and 2 grounded elements (the unaccounted-for grounded elements are the ones right at the 500k pot).
Regardless, if you sweep the 500k pot to one side (bottom), then you get a trebly sound, and if you sweep to the other side (top) you get a bassy sound.