Sometimes the least likely culprit is the culpable one after all. I had an amp one time that was a Marshally type with reverb (famous brand name). I always had problems with the reverb. I had tried everything in the book to try to figure it out to no avail.
The reverb worked fine, but it sure was sensative and at certain settings it would howl and also, it would cause hum problems.
I finally just resorted to only using it on settings that weren't problematic.
Months later, I decided that the amp would sound better if I changed it to use 6L6's instead of EL34's.
It not only sounded better with 6L6's, but the reverb problems vanished into thin air.
The point is, there are the usual suspects, and then there is everything else. Sometimes, you have to try everything.
Here, by the way, is my hunch on what the problem was...
With the EL34's, the power transformer (heater windings) were pushed pretty close to rated capacity. The power transformer itself would hum slightly when operated. With the change to 6L6's, it wasn't worked as hard and as a result, the humming stopped.
I think that while the trasformer was working so hard, it may have been producing more EMI and interfering with the reverb circuit.
Either that or the EL34's themselves were causing interference. Either way, the change to 6L6's was the fix.
Come to think of it, there were also problems with the built in EQ. If you set the bass too high, it would cause problems with annoying oscillations. That was fixed with the power tube change as well.
Dave