I still think the biggest problem is the speaker, or possibly tube choice. But probably speaker & cab.
I still do not understand why the plates are so high on the 12ax7s when all the resisters are correct and everything is wired proper.
Rather than worrying about why, let's just figure out how to "fix" it. 2ma of current might seem a little bit low for a 12AX7 but not unreasonable with 100K plate, 1.5K cathode, and 344 VDC high voltage supply for the preamp. However, 2ma is what you have (V/R=I so 20/10,000=2ma). FWIW I usually assume 1.5ma per 12AX7 triode for starters but 100K/1.5K rarely draws that much current.
Note that your B+ for the preamp is only 14 volts off spec - well within the +/- 20% allowance. For that matter, the 12AX7 plate voltages are only 15% above what's shown on the schematic and within spec.
If you replace the 10K dropping resistor on the power rail with a 27K resistor, you should get about 310 VDC supply for the 12AX7s (.002ma * 27,000 = 54 volts). That should drop your 12AX7 plate voltage(s) down below 200 volts and may give you a "rounder", mellower tone from the preamp.
You also could bias the first triode a bit hotter/differently. Either a 1K cathode resistor or a 168K to 200K plate resistor might be fun to try. Depends on how clean you want the amp to sound and how much headroom you want.
Another area to explore is the tone stack. Maybe a 330p treble cap would help. Tack a 100p cap in parallel with the 220p that's there now and listen. Maybe you'd like two .022uf mid/bass caps like a Marshall instead of Fender's .1uf bass & .047 mid.
For me, building an amp is like cooking. I start with a recipe, sure. I taste throughout the process and adjust proportions as I go along. Maybe I decide to leave something out (never the garlic though), maybe I throw something else in. Some things I measure. Most things I just eyeball in terms of quantity - I'm more interested in relative proportions. Maybe I make a note to do X differently next time. Everything from your finger tips to the speaker cone is part of the instrument, including guitar strings and coupling caps. That's a LOT of possible variables in combination. Too many variables to know exactly how it's going to turn out.
The AA764 is a classic circuit. If you have an NOS 12AX7, a Tung Sol 6V6 and a real 5Y3-GT, that's a start. Pine cab same dimensions plus a Weber 8A100-T speaker and you're on your way to an amp that should sound like a BF Champ would have in 1960 whatever.
The OT is a big part of any amp's sound too. Have you told us what the OT is? The voltage drop across the OT secondary is a little bit higher than "spec" which might indicate a higher secondary winding resistance. Whatever it is, the secondary winding resistance is greater than 1K. We know that because the voltage drop across the OT is greater than the voltage drop across the 1K dropping resistor going to the screen grid supply.
The point of all this is that you've built the amp to vintage specs (with the possible exception of the speaker/cab/OT). Voltages are within 20% of spec, so nothing's drastically wrong. You want it to sound different. Change it to taste the way
you like it.
Good luck,
Chip