> Might wanna leave the back end be although 4x 6AV5's is certainly making 18 watts the hard way
It's 75 Watts.
Power supply will run, full-load, 500-550V at 200-240mA.
OT primary is 4K-6K (probably near 5KCT); which is ~~10K per-pair.
What a wacky setup. Three small rectifier bottles. The absurd B+ has to be dropped way down so there is a TV V-sweep bottle making 200V ratio-ed (not regulated). Power section needs 1.5V-2V at V3A grid, so V2 must put out around 15V.
My gut says do as little as possible. You can play guitar into the MIC inputs. You might like it better changed over to 33K 1.5K+10u 100K Fender values. The volume and mixer network is already known-fine Fender values. Bogen output stages are usually musically benign.
For more pizzazz, increase R28 to 33K, 56K. Don't go a lot higher; this amp is high-strung and may hurt itself without NFB. Take the "Phono" (crystal) pot and put it between tone controls and V3A as Master; V2 sure can distort with just a little more loss after it.
R16 R27 look strange; they are semi-Akido networks. (There's nothing new under the sun.)
There's no obvious drop-in "guitar tube" output stage for this OT, even for this PT. If 6AV5 are really hard to get, we could consider pair-6550 (maybe quad-EL34), but they really should have G2 well below 550V more like 300V but the "200V" reg can't go any higher without stressing its heater insulation.