> the feedback wire ... What happens if you unsolder that...
He has a pot which will turn-down NFB to practically zero (<1:100, when forward gain is probably ~~20). Since that didn't fix the problem, I penciled-out bad NFB (my first guess also).
> one 1.5K resistor going to the grids of the output tubes. Before I had each tube with a resistor on it. Could that make a difference?
Yes, and circuit logic does not help.
You think each "side" of the push-pull quad will work together. Like a two-pair of oxen.
But it can be more like a two-pair of greyhounds. A pair which should "pull evenly", one dog gets a little ahead, then the other, very rapidly. Your V4 and V5, which are just "push" for audio, interact with the slightly different lead-lengths and stray capacitances so they act as a supersonic (radio frequency) push-pull oscillator. For a while. After a few thousand cycles the grid voltages has pounded itself negative, and the tubes cut-off. Either the cut-off or the recovery "pops!" This depends also on static voltage and grid resistance (including meter loading).
Yes, sometimes the layout and parts "get lucky" and it doesn't do that. (Until HBP puts in a slightly hotter tube set.)
Here's the conservative thing.
*Separate* resistors for each G1, G2, and plate.
In pentodes, the G1 resistor can be much larger than a K without real audio effect. 10K probably won't hurt. 5K is popular. Use separate 5K per G1.
G2 at 500 ohms is fine. Much smaller won't spoil supersonic troubles well, and much larger will restrict EL34 power.
Most good reliable quad-output amps have a small resistor in each *plate* lead. 47 ohms 2 Watts is common. This has hardly-measurable effect on audio power, but is enough to spoil the typically low-impedance radio resonances.
Yes, total of 12 resistors. You see why musician-market amplifiers try to get away with less. But DIY economics is different. You already spent more time head-banging than 12 resistors are worth.