Though a schematic for your Elk model seems hard find, here's one that seems to be a lower wattage version of your amp:
http://www.drtube.com/schematics/elk/es-30.gif It seems to have your preamp and PI tube tube line-up, except for the power tubes:
http://www.drtube.com/guitamp.htm So the schematic might be useful.
1. The trem circuit is likely identical. One-tube 12ax7 wobbling the tone recovery stage.
2. a) Pull tubes.
b) Disconnect the bias supply from the 2X 220K junction, at the bias end. What is the bias output voltage?
c) Pull the coupling caps; 220K's and grid stoppers (if any).
d) run the bias lead wire from the bias output to the the junction lug of the 220K's (no resistors present yet). What is the bias voltage at that point?
e) re-connect 1X 220K resistor at the bias end only - no connection yet on the power tube side. What is the bias voltage on the power tube side of this resistor?
f) same as e) for 2nd 220K resistor.
g) re-install 1X grid stopper. What is the bias voltage on the tube pin lug?
h) same as g) for 2nd grid stopper.
i) reconnect 1 coupling cap. What is the bias voltage on the tube pin lug?
j) same as i) for 2nd coupling cap.
Cathode bias. IMHO cathode bias is all about the power tube HT supply voltage. Specs say 375, but you've got more like 430. Unless you do a 50 volt drop (which is fine), your tubes will run hot in cathode bias. Cathode bias is dynamic, so that the tube always wants to draw the same idle current regardless of the size of the cathode resistor. E.g., because cathode bias is hot, you increase the size of the cathode resistor to limit current. But the larger cathode resistor pushes up the plate voltage on the other end of the circuit, which exerts a stronger pull on the cathode's electrons and re-establishes about the same current draw. And vice-versa. The only way to defeat this circle is to lower the B+ supply voltage to the plates & screens.