Sounds like you've checked the obvious and probable culprits.
Minor possibilities remaining;
Maybe the .003 output cap from the rev recovery stage is bad. (low possibility)
Are you sure your power supply caps are good? Especially the one/ones at the highest B+. The Rev driver gets big volts (compared to any of the other 12A_7's) and while you would expect such an issue to produce hum elsewhere, the output stage (the other place which gets the big B+) is in a push-pull config which tends to cancel hum. But that B+, applied to the single-ended reverb driver, could produce hum there which would pass into later amp stages. This line of thinking gets my biggest suspicion. Maybe your first 1 or 2 electrolytics are not doing their job. The preamp tubes have later, lower voltage / lower current electrolytics of their own and can probably tolerate
"less than normal" filtering from the first couple of e-caps. You could just try bridging the first or first & second e-cap(s) with 20 uf /450. Be careful, please!
You could also check for AC volts on that first and second filter caps. If you do, check both before and after you bridge in your temporary replacement caps.
And yes, it is absolutely possible that one cap in a multi-section electrolytic can go bad while others still work.