Or, if this is an already-built amp, just do the MOSFET regulator you drew to drop the voltage. You might have to adjust some resistor values on test to dial in the exact voltage you want.
... In order to drop 50 volts from the screen supply a mosfet will be used. Have heard mosfets prefer pulsating, unsmoothed voltage. Not sure why. Maybe the pulses give it a break? ...
I've heard a lot of things too. Usually, they are out of context.
MOSFET's only have so-high voltage ratings and so-high power dissipation ratings. If you have to drop a LOT of voltage, pulsating d.c. means the power dissipation in the MOSFET is less than if the voltage (and the implied resulting current, which is more important here) were unwavering. You're looking to drop 50v for screen which maybe hit 20mA peak (if that), times 4 tubes, which is 80mA * 50v = 4 watts. Even if I guessed only half-right, you can waste that much heat in a MOSFET properly insulated/bolted to the chassis.
If you use the MOSFET to filter/feed the output tubes, then it has to be rated for essentially all the current draw of the amp, times the voltage you wish to drop, plus some sensible safety margin. You could do this in the sense of an "overall power scale/VVR scheme" but I'll defer to others who know more about these circuits.