Oh and a good shot of the pots and jacks and the opposite side with the rca's and output connections, those aren't quite totally visible.
I'm not seeing anything specifically out of whack. I'd suggest getting voltage readings on eacy of the filter capacitor pins to ensure they're getting voltage and therefore are filtering correctly. Additionally, make sure to get voltage readings on all the important pins of every tube and post here. That will help ensure that the tubes are around the right voltage. Make sure you used the 40uF one for the first filter node, ( don't know from those caps for sure if that right top one is the 40uF, but make sure.) It could be motorboating going on.
Also do some thorough testing with continuity mode to check that continuity is working through every place it should.
The best way to do that is to go to some point before a turret, and then some point after it, don't measure right on the turret because you could be creating the connection by adding pressure on it.
Example, test continuity on the power rail by touching the leads of the 100k resistors between two nodes, that confirms you've got continuity between those leads, through the turret and through the jumper wires you've got there.
Check leads between resistors/capacitors and the tube pins by touching the lead of the component and the pin of the tube, not the turret, or wire itself.
Just be super anal about making sure EVERY connection is solid and has no bad connections.
Double check that your under board jumpers haven't somehow come undone in resoldering things, etc.
Check grounds from leads throughout the entire grounding busses, not on the turret or buss wire itself.
I'm not seeing anything that seems missing, but just thinking of other possible areas.
If you don't find anything, but give us the voltages it may point out a specific trouble area.
~Phil