Change tubes first. Then check voltages at all B+ nodes and ***ALL*** tube pins, even the pins that measure zero volts. Compare to voltages on this schematic/layout. Post voltages here if you need a hand. Signal tracing with a sig gen and a scope is a good idea too. Inject a 200mV 500Hz signal into the input jack and chase it through the amp.
I'm happy to have a hand for external verification if nothing else. All tubes replaced with known good as per tube chart except using 6L6GC rather than 6L6G. I'm also using a home made bucking transformer to knock the wall down to about 106VAC. Following voltages reflect this attenuated wall voltage. I see the grid of triode 2 of the phase inverter is significantly off (hopefully in red below)
V1 (12AY7)
plate 1: 125 plate 2: 123
grid 1 0 grid 2 0
cathode 2 cathode 2
V2 (12AY7)
plate 1: 129 plate 2: 126
grid 1 0 grid 2 0
cathode .001 cathode .001
V3 (12AY7)
plate 1: 111 plate 2: 226
grid 1 .01 grid 2 111
cathode 1.6 cathode 113.7
V4 (12AX7)
plate 1: 197 plate 2: 248.3
grid 1 .03
grid 2 18.3 cathode 1.3 cathode 46
V5, V6 (6L6GC)
Plates: 391
Grid 1: 393
Grid 2: -36
Cath -.001
V7, V8 (5U4GB): +394 on pins 8 and 2 (pins 1, 3, 5 not connected show .001 to .004; pins 4, 6 show "OL" DCV)
HEATERS
5.8 VAC
0 VDC except V4 (pins 4,5) and on V6 (pin 7) which goes between 0 and .001
*NOTE: there does appear to be a slight variation from the schematic in that the center tap of the OT has been moved upstream of the choke, the 6L6 suppressor grid voltage remains downstream