Hmmm, I wonder. Seems like the heaters may be dc sunken. Now there's even more voltage difference between cathode and filament. ...
Yikes!! That seems like a truly-bad idea, since there's no tube cathode running down to a negative rail.
... I think they did it for noise? ...
Can't be.
Heater-to-cathode leakage can induce hum via two mechanisms. One is to envision the heater-cathode as a diode that is forward-biased when the cathode is positive of the heater, thereby passing a 60Hz pulse of half-wave rectified hum to the cathode and into the audio.
That's why we usually have a positive voltage for elevating the heater: we're reverse-biasing the "heater-cathode diode" so that the cathode is never positive (relative to the heater) at any part of the heater's AC voltage cycle.
Tektronix (possibly others) would have a negative DC reference for heaters of some tubes. However, that was only for
some tubes, typically used as differential amplifiers whose cathode connected through 100-150kΩ resistors down to a -150v supply rail (the same tube might have 100-200kΩ up to a +350v supply). And there, it was mostly about reducing heater-to-cathode voltage (I'd have to look, but I think the reference was the -150v rail itself).