FWIW(1) I already have the 291AEX and would like to use that one. ... If (with zeners) I can get in the area of 300V on the plates ...
I can understand wanting to use a transformer that's already in-hand.
My preference is to use "the part that delivers the desired voltage" rather than use a kludge like zener to attempt to lose 100-150v.
... but won't the preamp lose clarity on lower voltages? ...
No. The biggest change is maximum AC output voltage that the preamp can deliver.
I have some outstanding vacuum tube mic preamps that only have a B+ of 250v. "Clarity" can be achieved (or discarded) with EQ, or by "indirect-EQ" in the form of output impedance based on your choice of operating point & plate load resistor.
... the 6AQ5 is merely an experiment. I drilled the chassis for 6V6 tubes and have the adapter plates for noval tubes installed (I'm using 6P1P/EL90 tubes, SED Winged=C=) If I don't like it I can always go to using 6V6 without a lot of hassle...
... You can get NOS SED =C= 6P1P for 5€. That's 5 to 6 times less than the price of an average current production 6V6.. so why not?
6P1P and 6AQ5 share a max voltage rating of 250v. So it's easier to build a safe environment with the lower-voltage transformer.
- 6V6 (with a Princeton's 8kΩ OT) delivers about 15 watts.
- If you lower the plate & screen to 250v (and use a 5k-6.6kΩ OT) you get 10w of output.
- 10 log (10w/15w) = -1.76 dB SPL
The output power reduction is so minor, you might miss it. OR, you will definitely fail to notice the power-reduction if you experience a "correct voltage" Princeton Reverb with an original-style Jensen C10Q or C10R, then use a Celestion or Eminence speaker for your "250v amp." The higher sensitivity of the latter speakers will make the 250v amp louder than the "higher-powered Princeton."