In your V1 pix, 2 posts above this one....I see a black heater feed wire going to pin 9 and I see a brown one going to pin 4.....and while I see there is solder filling up the hole in the lug on pin 5...I don't see a connection, a jumper if you will, between pin 4 and pin 5. It's perfectly possible that I am not seeing the jumper as it could be hiding under brown heater-feed wire. It just does not look like it's there. It's absolutely possible that if you are a newb, you may have wired the heater only to one lug. Or you wired it to 5, thought you made an error, then moved it to pin 4. Has to go to BOTH pins 4 & 5. Or there has to be a jumper between pins 4 & 5, however you wish to state it in words. Pins 4 & 5 are shorted together AND fed that brown wire.
If that jumper is not there, that tube will LOOK LIKE it is lit up, just quickly looking at the powered-up tube, but only half of the dual-triode will be lit up (you'll only see ONE dot of light atop the innards of the tube when looking at the tube powered up) but the unlit half will be dead as a doornail. I mean, DEAD, shut off, not working. As if it was pulled out of the amp.
My bet is that this is the problem. Turn the amp on and look at that V1 tube. No meters. Do you see two dots of light or just one? Compare to the other dudes.
So these 12A_7 triodes are two identical triodes or are supposed to be. Therefore, it does not matter if the very first triode your guitar signal hits is 1-2-3 or 6-7-8. The Hoffman board *probably* follows Fender but may not, I have not checked. It would not matter. But it *COULD* be that XYZ amp co and Hoffman designate the tube-halves differently.
All that is meaningless, you need not care. But you MUST care that 4 and 5 are to be jumpered together for the tube to be fully lit up if you are powering the heaters w/6.3 VAC, which you clearly are. If that tube has a dead (unlit) half, you better believe it could 100% kill your guitar signal.