Appears to be missing the artificial center tap as there is no filament center tap to ground. Confirm this with measuring the resistance of one leg to ground when powered off though. Some of these early 60s PTs have internally grounded filament CTs. Haven't seen it on a Twin Reverb though--but most of my experience with TRs is post '68...aka What my clients can afford.
The bias filter cap looks like it's several decades old. Its worth investigating. Put your meter in Vac and check if you have significant ripple riding on the bias voltage. You may have a 60Hz AC spike to low voltage that the DC measurements aren't picking up. (Grasping at straws with that one.)
The OT is definitely aftermarket, so that carries the risk of it being improper spec and installed incorrectly. You can try disconnecting the NFB at the speaker jack and rechecking bias to see if it was oscillating above audible frequencies which is causing the red plating. If this causes the bias to behave and the tubes to stop red plating, then reverse the secondaries and reconnect the NFB.
Finally, I always replace tubes that have a history of red plating. I would want a fresh set in there after you've fixed the problems, regardless. So perhaps have a set on order. This isn't as thoughtless a suggestion as it was even 10 years ago with the pricing being what it is now vs then. But in my experience, once they pop the fun don't stop--so a clean slate is warranted after the servicing is complete.